The answer choice D and E can be eliminated first because we are comparing same species to that of different species(not limited to one), and between can only be used when a comparision between A and B is made. 'Greater than' is an idiom and 'greater as' is not. Cancel A. Between B and C, the comparison is different species, plural, and in choice B, 'than that among individuals' makes it singular. 'Those' is plural and makes C the best answer.